Hyainailouros

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Hyainailouros
Temporal range: 20.0–11.3 Ma Early to Middle Miocene
Hyainailouros sulzeri.jpg
lower jaw of Hyainailouros sulzeri
Africa - the evolution of a continent and its large mammal fauna (2006) fig. 8.png
Restoration of H. sulzeri (far left), Cynelos eurydon, Afrosmilus africanus and H. napakensis (far right)
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Hyaenodonta
Superfamily: Hyainailouroidea
Family: Hyainailouridae
Subfamily: Hyainailourinae
Tribe: Hyainailourini
Genus: Hyainailouros
Biedermann, 1863
Type species
Hyainailouros sulzeri
Biedermann, 1863
Species
  • H. bugtiensis (Pilgrim, 1912)[1]
  • H. napakensis (Ginsburg, 1980)[2]
  • H. sulzeri (Biedermann, 1863)[3]
Synonyms
synonyms of genus:
  • Hainailouros (Lavrov, 1999)[4]
  • Hyaenaelurus (Stehlin, 1907)
  • Hyaenailurus (Rütimeyer, 1867)
synonyms of species:
  • H. napakensis:
    • Hyainailouros nyanzae (Ginsburg, 1980)
    • Pterodon nyanzae (Savage, 1965)[5]
  • H. sulzeri:
    • Hyainailouros maximus (Meyer, 1837)
    • Hyaenailurus sulzeri (Biedermann, 1863)

Hyainailouros ("hyena-cat") is an extinct polyphyletic genus of hyainailourid hyaenodont mammal of the polyphyletic tribe Hyainailourini within paraphyletic subfamily Hyainailourinae, that lived during the early to middle Miocene, of which there were at least three species spread across Europe, Africa, and Asia.[6][7]

Closely related to other large African hyaenodonts such as Simbakubwa and Megistotherium, Hyainailouros walked with a semi-digitigrade stance and was probably capable of large, leaping bounds.[8]

Alongside its African relatives and the last members of the genus Hyaenodon from Asia, Hyainailouros was among the largest hyaenodonts that existed.[citation needed]

Phylogeny

The phylogenetic relationships of genus Hyainailouros are shown in the following cladogram:[9][8][10][11]

 †Hyaenodonta 

Eoproviverra

Boualitomidae

Arfiidae

Limnocyonidae

Sinopidae

Hyaenodontoidea

Gazinocyon

Pyrocyon

Galecyon

 †Afro‑Arabian clade 

Indohyaenodontidae

Glibzegdouia

Parvavorodon

Koholiidae

Tritemnodon

Furodon

Kyawdawia

Paratritemnodon

Teratodontidae

Hyainailouroidea

Apterodontinae

Orienspterodon

Hyainailourinae

Hemipsalodon

 ? 

Ischnognathus

Akhnatenavus clade

Akhnatenavus

Paroxyaenini

"Pterodon" sp. (BC 15’08)

Hyainailourinae sp. (UON 84-359)

Hyainailourinae sp. C (DPC 9243 & DPC 10315)

Hyainailourinae sp. D (DPC 6545)

Hyainailourinae sp. A (DPC 6555)

"Pterodon" africanus

Parapterodon

"Pterodon" sp. (DPC 5036)

"Pterodon" phiomensis

Hyainailourini

Kerberos

"Pterodon" syrtos

Pterodon clade

Pterodontina

Falcatodon

Sectisodon

Exiguodon

Isohyaenodon zadoki

Isohyaenodon  (†Isohyaenodontina)

Isohyaenodon andrewsi

Sivapterodon

Hyainailouros bugtiensis

Hyainailouros napakensis

Hyainailouros

Hyainailouros sulzeri

Hyainailourinae sp. (GSN AD 100’96)

Simbakubwa

Leakitheriini

Megistotherium

Mlanyama

Metapterodontini

Pakakali

Prionogalidae

Lahimia clade
Arfia clade
Galecyon clade
Indohyaenodon clade
Tritemnodon clade
Kyawdawia clade

References

  1. ^ Pilgrim, G. E. (1912.) "The Vertebrate Fauna of the Gaj Series in the Bugti Hills and the Punjab", Memoir of the Geological Survey of India, Palaeontologia Indica, New Series, 4: 1–83
  2. ^ Ginsburg, L. (1980.) "Hyainailouros sulzeri, mammifère créodonte du Miocène européen." Annales de Paléontologie, 66: 19–73.
  3. ^ W. G. A. Biedermann (1863.) "Petrefacten aus der Umgegend von Winterthur. II Heft: Die Braunkohlen von Elgg. Anhang: Hyainailouros sulzeri". Bleuler-Hausheer, Winterthur, 23 pp.
  4. ^ A. V. Lavrov (1999.) "Adaptive Radiation of Hyaenodontinae (Creodonta, Hyaenodontidae) of Asia." in 6th Congress of the Theriological Society, Moscow, April 13–16, p. 138 [in Russian].
  5. ^ R. J. G. Savage (1965.) "Fossil Mammals of Africa: The Miocene Carnivora of East Africa." Bulletin of the British Museum (Natural History) Geology 10(8):241-316
  6. ^ Morlo, M.; Miller, E. R.; El-Barkooky, A. N. (2007). "Creodonta and Carnivora from Wadi Moghra, Egypt". Journal of Vertebrate Paleontology. 27: 145–159. doi:10.1671/0272-4634(2007)27[145:CACFWM]2.0.CO;2.
  7. ^ Solé, F.; Lhuillier, J.; Adaci, M.; Bensalah, M.; Mahboubi, M.; Tabuce, R. (2013). "The hyaenodontidans from the Gour Lazib area (?Early Eocene, Algeria): implications concerning the systematics and the origin of the Hyainailourinae and Teratodontinae". Journal of Systematic Palaeontology. 12 (3): 303–322. doi:10.1080/14772019.2013.795196.
  8. ^ a b Borths, M. R.; Stevens, N. J. (2019). "Simbakubwa kutokaafrika, gen. et sp. nov. (Hyainailourinae, Hyaenodonta, 'Creodonta,' Mammalia), a gigantic carnivore from the earliest Miocene of Kenya". Journal of Vertebrate Paleontology. 39: e1570222. doi:10.1080/02724634.2019.1570222. S2CID 145972918.
  9. ^ Borths, Matthew R.; Stevens, Nancy J. (2017). "Deciduous dentition and dental eruption of Hyainailouroidea (Hyaenodonta, "Creodonta," Placentalia, Mammalia)". Palaeontologia Electronica. 20 (3): 55A. doi:10.26879/776.
  10. ^ Floréal Solé; Bernard Marandat; Fabrice Lihoreau (2020). "The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian". Geodiversitas. 42 (13): 185–214. doi:10.5252/geodiversitas2020v42a13.
  11. ^ Solé, F.; Morlo, M.; Schaal, T.; Lehmann, T. (2021). "New hyaenodonts (Mammalia) from the late Ypresian locality of Prémontré (France) support a radiation of the hyaenodonts in Europe already at the end of the early Eocene". Geobios. in press. doi:10.1016/j.geobios.2021.02.004.

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