Manis palaeojavanica

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Manis palaeojavanica
Temporal range: 0.8–0.03 Ma
late Pleistocene
Manis palaeojavanica and javanica.jpg
bones of Manis palaeojavanica (darker)
compared to same bones of
Manis javanica (lighter)
Scientific classification edit
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Pholidota
Family: Manidae
Genus: Manis
Subgenus: Paramanis
Species:
M. palaeojavanica
Binomial name
Manis palaeojavanica
Dubois, 1907[1]
Synonyms
list of synonyms:
  • Manis palaejavanica (Dubois, 1908)[2]
  • Manis palaejavanico (Weber, 1928)[3]
  • Manis palaeojavanicus (Dubois, 1907)
  • Manis paleojavanica (Raven, 1935)[4]

Giant asian pangolin (Manis palaeojavanica ["ancient Javan pangolin"]), is an extinct species of pangolin (from genus Manis) that was native to Asia.[5]

In 1926, E. Dubois described the bones of M. palaeojavanica discovered in Java. Later, Lord Medway excavated another set of bones at the Niah Caves in Malaysia. In 1960, D. A. Hoojier identified these bones as that of an extinct species. Using carbon dating, the Niah Caves bones were determined to be 42,000–47,000 years old.[6]

M. palaeojavanica is one of the only known fossils to be found nearly complete of the pangolin species, due to their armour-like scales that protected their bodies when they were alive.[7] Its total length is measured up to 2.5 m (8.2 ft).[8]

Phylogeny

Phylogenetic position of Manis palaeojavanica within family Manidae.[9][10][11]

 Pholidotamorpha 

Palaeanodonta Metacheiromys DB152-2.jpg

 Pholidota 

Euromanis

 ? 

Pholidota sp. (BC 16’08)

Eurotamanduidae

 Eupholidota 

Eomanoidea

 Manoidea 

Patriomanidae

 ? 

Necromanis

 Manidae 
 Smutsiinae 

Phatagininae Anatomische Untersuchungen über die Edentaten (1852) Phataginus tricuspis.png

Smutsiinae (sensu stricto) Cambridge Natural History Mammalia Fig 109.jpg

 sensu lato 
 ? 

Manidае sp. (DPC 3972 & DPC 4364)

 Maninae 
 Manis 

Manis pentadactyla

 ? 

Manis hungarica

Manis crassicaudata Pangolin Hardwicke (white background).jpg

 ? 

Manis lydekkeri

 ? 

Manis sp. (Scale_H4 & Scale_H8)

 (Paramanis

Manis culionensis

Manis javanica Anatomische Untersuchungen über die Edentaten (1852) Manis javanica.jpg

Manis palaeojavanica

 sensu stricto 
 (Pholidota sensu lato) 

References

  1. ^ Dubois, E. (1907.) "Eenige van Nederlandschen kant verkregen uitkomsten met betrekking tot de kennis der Kendeng-fauna (Fauna van Trinil)." Tijdschr. Kon. Ned. Aardr. Gen., Serie 2 (24):449-458
  2. ^ Dubois, E. (1908.) "Das Geologische Alter der Kendengoder Trinil-Fauna." Tijdschr. Kon. Ned. Aardr. Gen., Serie 2 (25):1235-1270, pl. 39.
  3. ^ Max Weber (1928.) "Die Säugetiere", vol. 2, Jena, p. 185
  4. ^ Raven, Henry Cushier (1935.) "Wallace's line and the distribution of Indo-Australian mammals." Bulletin of the AMNH; v. 68, article 4
  5. ^ John C. Barentine (2016). Uncharted Constellations: Asterisms, Single-Source and Rebrands. Springer. p. 186. ISBN 978-3-319-27619-9.
  6. ^ Quentin Phillipps; Karen Phillipps (2016). Phillipps' Field Guide to the Mammals of Borneo and Their Ecology: Sabah, Sarawak, Brunei, and Kalimantan. Princeton University Press. p. 153. ISBN 978-0-691-16941-5.
  7. ^ Gaudin, Timothy J.; Emry, Robert J.; Wible, John R. (December 2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis". Journal of Mammalian Evolution. 16 (4): 235–305. doi:10.1007/s10914-009-9119-9. ISSN 1064-7554. S2CID 1773698.
  8. ^ Hooijer, D.A., 1947 - A femur of Manis palaeojavanica Dubois from Western Java - Proceedings of the Koninklijke Nederlandsche Akademie van Wetenschappen 50 (4): 423-418
  9. ^ Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. Heidelberg, Germany: Springer Science+Business Media. 16 (4): 235–305. doi:10.1007/s10914-009-9119-9. S2CID 1773698.
  10. ^ Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. doi:10.1080/02724634.2012.694319. ISSN 0272-4634. S2CID 86059673.
  11. ^ Philippe Gaubert, Agostinho Antunes, Hao Meng, Lin Miao, Stéphane Peigné, Fabienne Justy, Flobert Njiokou, Sylvain Dufour, Emmanuel Danquah, Jayanthi Alahakoon, Erik Verheyen, William T Stanley, Stephen J O’Brien, Warren E Johnson, Shu-Jin Luo (2018) "The Complete Phylogeny of Pangolins: Scaling Up Resources for the Molecular Tracing of the Most Trafficked Mammals on Earth" Journal of Heredity, Volume 109, Issue 4, Pages 347–359

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