Achatinidae

Achatinidae; Temporal range: Eocene–Recent PreꞒ Ꞓ O S D C P T J K Pg N
	Giant East African Snail, Lissachatina fulica
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Heterobranchia
Order:	Stylommatophora
Suborder:	Achatinina
Superfamily:	Achatinoidea
Family:	Achatinidae; Swainson, 1840
Diversity
	176 extant species and subspecies in 16 genera within Achatinidae sensu stricto

Achatinidae (New Latin, from Greek "agate") is a family of medium to large sized tropical land snails, terrestrial pulmonate gastropod mollusks from Africa.

Well known species include Achatina achatina the Giant African Snail, and Lissachatina fulica the Giant East African Snail.

As of 2011 there were recognized 176 extant species and subspecies in 16 genera within Achatinidae.^[2]

Distribution[edit]

The native distribution of Achatinidae is Africa south of the Sahara.^[3]

Anatomy[edit]

In this family, the number of haploid chromosomes lies between 26 and 30 (according to the values in this table).^[4]

Taxonomy[edit]

The family Achatinidae is classified within the informal group Sigmurethra, itself belonging to the clade Stylommatophora within the clade Eupulmonata (according to taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).^[5]

As of 2017, the family Achatinidae contains the following subfamilies:^[6]

Achatininae Swainson, 1840 - synonyms: Urceidae Chaper, 1884; Ampullidae Winckworth, 1945
Coeliaxinae Pilsbry, 1907
Cryptelasminae Germain, 1916
Glessulinae Godwin-Austin, 1920
Opeatinae Thiele, 1931
Petriolinae Schileyko, 1999
Pyrgininae Germain, 1916
Rishetiinae Schileyko, 1999
Rumininae Wenz, 1923
Stenogyrinae P. Fischer & Crosse, 1877
Subulininae P. Fischer & Crosse, 1877
Thyrophorellinae Girard, 1895

Genera[edit]

Burtoa nilotica shell

Genera in the family Achatinidae include:^[6]

Achatininae

Achatina Lamarck, 1799 - type genus of the family Achatinidae^[5]
Archachatina Albers, 1850^[7]
Atopocochlis Crosse & P. Fischer, 1888
Bequaertina Mead, 1994
Brownisca Mead, 2004
Bruggenina Mead, 2004^[8]
Burtoa Bourguignat, 1889
Callistoplepa Ancey, 1888
Cochlitoma Férussac, 1821
Columna Perry, 1811
Leptocala Ancey, 1888
Leptocalina Bequaert, 1950
Leptocallista Pilsbry, 1904
Lignus Gray, 1834
Limicolaria Schumacher, 1817
Limicolariopsis d'Ailly, 1910
Lissachatina Bequaert, 1950
Metachatina Pilsbry, 1904
Pseudachatina Albers, 1850

Coeliaxinae

Balfouria Crosse, 1884
Coeliaxis H. Adams & Angas, 1865
Ischnocion Pilsbry, 1907
Nannobeliscus Weyrauch, 1967
Neosubulina Smith, 1898
Riebeckia von Martens, 1883

Cryptelasminae

Cryptelasmus Pilsbry, 1907
Thomea Girard, 1893

Glessulinae

Glessula Martens, 1860

Opeatinae

Eremopeas Pilsbry, 1906
Opeas Albers, 1850

Petriolinae

Bocageia Girard, 1893
Ceras Dupuis & Putzeys, 1901
Chilonopsis Fischer de Waldheim, 1848
Cleostyla Dall, 1896
Comoropeas Pilsbry, 1906
Dictyoglessula Pilsbry, 1919
Homorus Albers, 1850
Ischnoglessula Pilsbry, 1919
Itiopiana Preston, 1910
Kempioconcha Preston, 1913
Liobocageia Pilsbry, 1919
Mabilliella Ancey, 1886
Nothapalinus Connolly, 1923
Nothapalus von Martens, 1897
Oleata Ortiz de Zarate, 1959
Oreohomorus Pilsbry, 1919
Petriola Dall, 1905
Subuliniscus Pilsbry, 1919
Subulona Martens, 1889

Pyrgininae

Pseudobalea Shuttleworth, 1854
Pyrgina Greef, 1882

Rishetiinae

Bacillum Theobald, 1840
Eutomopeas Pilsbry, 1946
Rishetia Godwin-Austen, 1920
Tortaxis Pilsbry, 1906

Rumininae

Krapfiella Preston, 1911
Lubricetta Haas, 1928
Namibiella Zilch, 1954
Rumina Risso, 1826
Xerocerastus Kobelt & Möllendorff, 1902

Stenogyrinae

Chryserpes Pilsbry, 1906
Cupulella Aguayo & Jaume, 1948
Dolicholestes Pilsbry, 1906
Lyobasis Pilsbry, 1903
Neobeliscus Pilsbry, 1896
Obeliscus Beck, 1837
Ochroderma Ancey, 1885
Ochrodermatina Thiele, 1931
Ochrodermella Pilsbry, 1907
Plicaxis Sykes, 1903
Promoussonius Pilsbry, 1906
Protobeliscus Pilsbry, 1906
Rhodea Adams & Adams, 1855
Stenogyra Shuttleworth, 1854
Synapterpes Pilsbry, 1896
Zoniferella Pilsbry, 1906

Subulininae

Allopeas H. B. Baker, 1935
Beckianum Baker, 1961
Curvella Chaper, 1885
Dysopeas Baker, 1927
Euonyma Melvill & Ponsonby, 1896
Hypolysia Melvill & Ponsonby, 1901
Lamellaxis Strebel & Pfeffer, 1882
Leptinaria Beck, 1837
Leptopeas Baker, 1927
Micropeas Connolly, 1923
Neoglessula Pilsbry, 1909
Paropeas Pilsbry, 1906
Pelatrinia Pilsbry, 1907
Prosopeas Mörch, 1876
Pseudoglessula Boettger, 1892
Pseudopeas Putzeys, 1899
Striosubulina Thiele, 1933
Subulina Beck, 1837
Vegrandinia Salvador, Cunha & Simone, 2013
Zootecus Westerlund, 1887

Thyrophorellinae

Thyrophorella Greeff, 1882

References[edit]

^ Swainson W. J. (1840). A treatise on malacology161: 334.
^ ^a ^b Protiva T. (2011). Oblovky plži čeledi Achatinidae. Robimaus, 72 pp., ISBN 9788087293225.
^ "Family summary for Achatinidae". AnimalBase, last change 24-11-2007, accessed 3 August 2010
^ Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
^ ^a ^b Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
^ ^a ^b Bank, R. (2017). Classification of the Recent terrestrial Gastropoda of the World. Last update: July 16th, 2017.. World Register of Marine Species, Retrieved 10 January 2019.
^ Achatininae in www.petsnails.co.uk
^ Mead A. R. (2004). "Comparative reproductive anatomy in the South African giant land snails (Gastropoda: Pulmonata: Achatinidae)". Zoologische Mededelingen Leiden 78: 417-449.

External links[edit]

Data related to Achatinidae at Wikispecies
Media related to Achatinidae at Wikimedia Commons
Achatinidae
Extensive information on Achatina and Archachatina in captivity with a helpful forum.
US Dept of AgricultureError: "Q386510" is not a valid Wikidata entity ID.

[1] Swainson W. J. (1840). A treatise on malacology161: 334.

[Protiva_2011-2] Protiva T. (2011). Oblovky plži čeledi Achatinidae. Robimaus, 72 pp., ISBN 9788087293225.

[AB-3] "Family summary for Achatinidae". AnimalBase, last change 24-11-2007, accessed 3 August 2010

[Barker-4] Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.

[Bouchet_2005-5] Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.

[bank-6] Bank, R. (2017). Classification of the Recent terrestrial Gastropoda of the World. Last update: July 16th, 2017.. World Register of Marine Species, Retrieved 10 January 2019.

[7] Achatininae in www.petsnails.co.uk

[8] Mead A. R. (2004). "Comparative reproductive anatomy in the South African giant land snails (Gastropoda: Pulmonata: Achatinidae)". Zoologische Mededelingen Leiden 78: 417-449.

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