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Exmoor pony

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Exmoor pony
ExmoorponyPose.jpg
A herd of Exmoor ponies
Country of originUnited Kingdom
Traits
Distinguishing featuresMealy markings around eyes and muzzle, "snow chute/ice tail", "toad eye"

The Exmoor pony is one of the mountain and moorland pony breeds native to the British Isles. Some still roam as semi-feral livestock on Exmoor, a large area of moorland in Devon and Somerset in southwest England. The Exmoor has been given "Priority" conservation status by the Rare Breeds Survival Trust (UK), and "threatened" status by The Livestock Conservancy (USA). Exmoor ponies are particularly well-adapted to cold and wet weather and therefore very hardy, having conformation similar to that of other cold-weather-adapted pony breeds. The free-roaming herds on Exmoor contribute to the conservation and management of several natural habitats. Most foals born into the free-living herds are sold and with taming and training undertake a variety of equestrian activities.

Equines have been present in Britain since 700,000 BC, with populations retreating south when ice-ages made the land uninhabitable and recolonising when warmer conditions allowed. Some theorise that Exmoor ponies have changed very little from these prehistoric ponies from which they are descended: others consider them as simply one of the relatively modern created breeds of pony. In morphological and appearance (phenotypically) Exmoors closely resemble the primitive wild horse and modern DNA research is trying to determine exactly how much of the ancient genome, if any, survives in the Exmoor pony.

Written records of ponies on Exmoor start with the Domesday Book in 1086. After centuries of being a “Royal Forest” (not an area of trees but a hunting ground), most of Exmoor was sold in 1818. Thirty ponies, identified as the original old type, were saved and relocated to neighbouring moorland. These ponies form the foundation bloodstock of today's Exmoor population. The Exmoor pony breed society was formed in 1921 as the Exmoor Pony Society.

The breed nearly became extinct during the Second World War owing to ponies being stolen for the wartime meat market and a few being used for target practice by soldiers. After the war a small group of breeders worked to save the Exmoor, and during the 1950s ponies began to be exported to continental Europe and to Canada in small numbers. The first stud book was published in 1963 and numbers grew slowly. In 1981 publicity regarding the vulnerability of the breeding population raised interest in the need to conserve the Exmoor pony and its numerical recovery (total population size) gathered pace. By 2021, 100 years on from the formation of the Exmoor Pony Society, approximately 600 ponies roam Exmoor and a further 4,000 (some free-living but most as domestic ponies) are in locations across the UK and in several other countries. The vast majority are in non-breeding situations, hence the RBST’s “Priority” classification.

Characteristics

An Exmoor mare and foal

The Exmoor pony coat colour is predominantly a variant of dark bay, called "brown", with some individuals lighter and termed "bay" and rarely dun coloured. All Exmoors have to display pangaré ("mealy") markings[1] around the eyes, muzzle and underbell to be registered. Black has been bread out of the Exmoor breed because pangaré does not show on black based horses. Pangaré markings occur in other equines as well as horses, and are considered to be a primitive trait. In order to be registered in the stud book Exmoor ponies cannot have white markings. There is at least one ancient herd with red dun exmoors in England and several herds exist in which bay dun exmoors are present. Dun, bay and seal brown are acceptable in exmoors when being registerd. Sooty (countershading) is common in exmoors. Most exmoor ponies are homozygotic for the non-dun 2 gene, which is a recessive gene that developed after domestication and causes a lack of dilution and an absence of wild type markings. In the exmoor breed there has no evidence been found yet for the occurence of the wild type non-dun 1 gene, which causes non-dilution or limited dilution with some wild type markings visible (most commonly a dorsal stripe). In height, the majority of Exmoors range from 11.1 to 12.3 hands (45 to 51 inches, 114 to 130 cm) (45 to 51 inches, 114 to 130 cm), with the recommended height limit for mares being 12.2 hands (50 inches, 127 cm) (50 inches, 127 cms) and that for stallions and geldings 12.3 hands (51 inches, 130 cm). A few reach 13.2 hands [2]

With a stocky, powerful build, the Exmoor pony is strong for its height, and noted for its hardiness and endurance.[3] The chest is deep and the back broad, the croup level. The legs are short, with good bone and hard hooves.[2] Although many sources state that the Exmoor has a distinctly different jaw structure from other horse breeds, which includes the beginnings of development of a seventh molar, this is a misunderstanding based on an incorrect translation of a German study.[4] The study refers to an extra branch of blood supply to the jaw and speculates that this might have been the early stages of evolving an extra molar. However the feature is found in a number of horse breeds with sizable lower jaws, so its presence in the Exmoor pony is unremarkable.[5][6]

The head is somewhat large in proportion to the body, with small ears, and has a special adaptation called a "toad eye" (raised flesh above and below the eyes) which helps to deflect water and provide extra insulation. As with most cold-weather adapted pony breeds, the Exmoor grows a winter coat consisting of a highly insulating woolly underlayer and a topcoat of longer, oily hairs that prevent the undercoat from becoming waterlogged by diverting water down the sides of the animal to fall from just a few drip areas. The mane and tail are thick and long, and the dock of the tail has a fan of shorter, coarse hairs, called a "snow chute" that deflects rainwater away from the groin and underbelly areas to fall from the long hairs on the back of the hind legs.[3][5]

History

A group of Exmoor ponies
A herd of Exmoor ponies

Ponies were first recorded in the Exmoor area in the Domesday Book in 1086, but no count of the ponies roaming the Royal Forest of Exmoor is included as, being owned by the King, they were not to be taxed. Over the centuries, a series of Wardens managed the Royal Forest, charging fees for the grazing of livestock including ponies. The change of Warden was often accompanied by the sale of ponies they personally owned, as for example in 1748 at the end of the Hill and Darch wardenship.[7] Whether some were sold to incoming Wardens is not recorded. In 1767 Warden Sir Thomas Acland was grazing ponies on the Forest. Sales records from 1805 and 1809 list the colours black, grey, bay, dun, 'chasnut', and piebald for ponies. Studies disagree as to the significance of this: some argue it shows that the indigenous brown and mealy type of Exmoor pony had probably been lost through cross-breeding;[8] others consider that because only the Warden could run stallions, cross-bred mares of different colours could co-exist with the indigenous type.[9]

In 1818 the Royal Forest and some adjoining land was sold to John Knight. Sir Thomas Acland, the outgoing warden, took thirty of “the old type” to Ashway Side close to Winsford Hill, while others were sold to local farmers. Records show that Acland then adopted two breeding strategies, a closed, purebred herd on Ashway Side and some separate crossing experiments on Winsford Hill.[10] In contrast, John Knight, starting in 1826, crossed all the ponies he had purchased with Arabians, Thoroughbreds, and other breeds to increase their size.[11] Significantly, today’s registered Exmoor pony population is descended from the Acland herd.

From the late nineteenth century, Exmoors were selectively bred for the characteristic mealy muzzle. In 1921, the fledgling Exmoor Pony Society’s rules accepted grey and black ponies in addition to bay, brown and dun, though the colour standards were soon tightened to exclude animals that did not conform to the carefully defined breed standard (based upon “the old type”).[6][9] Exmoor pony registrations (as well as any ponies from Exmoor) were recorded in the Polo & Riding Pony Stud Books (the National Pony Society Stud Books from 1913). The Exmoor Pony Society published its first stud book exclusively for registered Exmoor ponies in 1963.[12][11]

The Second World War led to a sharp decrease in the breed population. This was largely due to ponies being stolen for the wartime horse-meat market; a small number of ponies were supposedly also used for target practice by soldiers, Exmoor being a training ground.[6][10]

After the war, a small group of breeders, including Mary Etherington,[13] worked to restore the herds. In the 1950s, Exmoors were first exported to Canada and continental Europe, where small populations are still maintained. [14] Since the mid-1980s, small free-living herds of Exmoor ponies have been established in various areas of the UK: these herds are used to manage vegetation on nature reserves belonging to organisations such as the National Trust, Natural England, and County Wildlife Trusts.[15] Since the start of the 21st century Exmoor ponies have been in demand for ‘rewilding’ projects both UK-based and in additional European countries.

In 2000, the Moorland Mousie Trust, a British organisation, was established to assist in the preservation of the Exmoor pony. With very limited demand for Exmoor colts, the organisation works to raise funds for the gelding, handling and training of these ponies. In 2006, the Trust opened the Exmoor Pony Centre, on Exmoor.

Until 2009, every purebred registered Exmoor was hot branded for individual identification. Then, with microchipping offering a viable alternative, in-ground (not moorland herds) breeders ceased to brand. A few years later, hot branding became illegal throughout Britain for all except semi-feral ponies, for which there is no other workable alternative.

Three small brown horses on grassy area of Exmoor. In the distance are hills.
Exmoor ponies in their native habitat

Currently, the UK Rare Breeds Survival Trust watchlist includes the Exmoor breed in its most threatened category, "Priority", based upon the number of breeding animals and the level of in-breeding. Whilst worldwide the total population numbers around 4,000 (the vast majority within the UK), Exmoor Pony Society estimates the number of breeding animals is only around 600. The Livestock Conservancy in North America also considers the population of the Exmoor pony to be at "threatened" levels.[16]

Prehistoric origin theories

See also: History of the horse in Britain

Some claim that the Exmoor pony descends directly from the wild horses of north-west Europe, uninfluenced by domestic horses.[17] Others consider that Exmoor ponies have a fully domesticated origin.[6] Modern DNA research has not yet conclusively determined the breed’s origins: studies indicate they share their maternally inherited mitochondrial DNA with various other horse breeds from across the world,[18] and their paternally inherited Y-chromosome is identical to that of most other domesticated horses.[19]

Wild horses have lived in Britain for hundreds of thousands of years. Some remains found date as early as 700,000 BC,[20] while others are as recent as 3,500 BC.[21] No genetic studies to date have correlated these prehistoric remains to any modern breed. What has been studied are Y-chromosomes (Y-DNA) and mitochondrial DNA (mtDNA). The Y-chromosome is passed on through the male line, and worldwide shows very little genetic variation in horses,[19] except for a second Y-chromosome haplotype found in China,[22] suggesting that a very limited number of stallions contributed to the original genome of the domestic horse. The Exmoor pony shares this general Y-chromosome haplotype.[19] In contrast, mitochondrial DNA is passed on though the female line, and shows far more variation than Y-DNA, indicating that a large number of wild mares from several regions have contributed to modern domestic breeds.[23][24] Some mtDNA-haplotypes have been found in DNA samples obtained from wild horses in prehistoric deposits, while other mtDNA-haplotypes have only been found in domesticated horses, from both living individuals and archeological finds.[18] The Exmoor pony has a high frequency of pre-domestication mtDNA-haplotypes; these are also found in some other breeds all around the world.[18] Currently, for the British Isles, few DNA archeological samples have been studied.[25]

Although wild horses were abundant after the last ice age,[26] the lack of sufficient pre-domestication DNA samples makes it impossible to determine the contribution of the wild horses of the British Islands to modern breeds, including the Exmoor pony.[25] A 1995 study of morphological characteristics, the outward appearance of organisms, indicated that the Exmoor, the Pottock, and the now-extinct Tarpan have an extremely close resemblance; these breeds were consistently grouped together in the results from several analyses, with the Exmoor showing the closest relationship to the Tarpan of all the breeds studied, at 0.27; the next-closest breeds to the Tarpan were the Pottock and Merens, both with a genetic distance from the Tarpan of 0.47. The distance between the Exmoor and Pottock was 0.37, and between the Exmoor and Merens was 0.40; a significantly wider gap than the distance between the Exmoor and Tarpan.[27]

The first indication of domesticated horses in England comes from archaeological investigations showing that the ancient Britons were using wheeled horse-drawn transport extensively in southwest England as early as 400 BC.[28] Recent research has indicated that there was significant Roman involvement in mining on Exmoor.[29] Metals including iron, tin, and copper were transported to Hengistbury Head in neighbouring Dorset for export,[30] and Roman carvings, showing British and Roman chariots pulled by ponies phenotypically similar to the Exmoor, have been found in Somerset.[31][32][33][34]

Uses

Prior to the arrival of agricultural mechanisation, Exmoor ponies removed from the moorland herds, tamed and trained, were used primarily in hill farming. Tasks undertaken were ploughing/harrowing, shepherding and transport of feed. They were both ridden and driven, providing transport for Exmoor inhabitants. Some were sold away from the moor to work in harness elsewhere, possibly a small number as pit ponies.[35]

Since these past roles ceased, surplus moorland-bred foals and ponies bred in-ground (on farms, studs etc.) are used for a variety of leisure activities including showing, long-distance riding, driving,[3] and agility. In 2003 Stowbrook Jenny Wren was the first Exmoor to win the NPS Bailey's Horse Feeds Ridden Mountain & Moorland Championship at the Olympia Christmas Show; Exmoor ponies won both divisions at the International Horse Agility Championships in 2011,[36] and took a third title at the 2012 Championships.[37] The breed's hardiness, dentition and varied diet makes it suitable for conservation grazing, and it contributes to the management of many heathland, chalk grassland and other natural pasture habitats, as well as to the conservation of Exmoor itself.[15]

In January 2015, fourteen Exmoor ponies, from breeders within Exmoor National Park were exported to the Czech Republic's former military base of Milovice, to improve biodiversity by conservation grazing. According to the annual count of 2019, this population has increased to 111 ponies. [38] [39]

Gallery

  • Bampton Fair- Pony, Sheep and Cattle Sales in Bampton, Devon, UK, October 1943. Ponies were often sold here after gathering in the autumn.

    Bampton Fair- Pony, Sheep and Cattle Sales in Bampton, Devon, UK, October 1943. Ponies were often sold here after gathering in the autumn.

  • Exmoor Stallion Dunkery Wigeon at the Royal Highland Show 2018

    Exmoor Stallion Dunkery Wigeon at the Royal Highland Show 2018

References

Notes

  1. ^ Sponenberg 2003, p. 30.
  2. ^ a b "Rules of the Exmoor Pony Society — Breed Standard". Exmoor Pony Society. May 2013. Archived from the original on 19 April 2014. Retrieved 18 April 2014.
  3. ^ a b c Dutson 2005, p. 301.
  4. ^ Ebhardt, H (1962). "Ponies und Pferde im Rontgenbild nebst einigen stammesgeschichtlichen Bemerkungen dazu". Saugetierkundliche Mitteilungen. BLV Verlagsgesellschaft mbH.
  5. ^ a b "Exmoor Pony". International Museum of the Horse. Retrieved 15 April 2014.
  6. ^ a b c d Baker 2008.
  7. ^ Burton, Roger (1989). The Heritage of Exmoor. R. A. Burton. p. 55. ISBN 978-0951441909.
  8. ^ Green, Peter. "The free-living ponies within the Exmoor National Park: their status, welfare and future" (PDF). Exmoor National Park. Retrieved 13 November 2020.
  9. ^ a b Baker, Sue (2017). Sir Thomas Acland – did he really conserve the true Exmoor pony?. Exmoor Pony Society.
  10. ^ a b Orwin, C. S. (1970). The reclamation of Exmoor Forest. Newton Abbot: David & Charles. p. 69. ISBN 978-0-7153-4959-5. OCLC 130702.
  11. ^ a b Dent & Hendricks 2007, pp. 180–181.
  12. ^ "Exmoor Ponies – a dying breed?". BBC Somerset. Retrieved 3 December 2007.
  13. ^ "Persecuted Exmoor ponies trot back from the brink". Retrieved 9 October 2018.
  14. ^ "Exmoor Pony". Oklahoma State University. Retrieved 2 December 2010.
  15. ^ a b "Map of UK Conservation Grazing Schemes". Grazing Animals Project. 18 April 2012. Archived from the original on 14 March 2014. Retrieved 15 May 2012.
    • "Wildlife Conservation of Local Downland and Heathland". Sussex Pony Grazing and Conservation Trust. Retrieved 15 May 2012.
    • "Grazing Exmoor ponies to protect County Durham flowers". BBC News. 8 March 2011. Retrieved 15 May 2012.
  16. ^ "Breed Information – ALBC Conservation Priority List". American Livestock Breeds Conservancy. Retrieved 2 December 2010.
  17. ^ Hovens, Hans; Rijkers, Toon. "On the origins of the Exmoor pony: did the wild horse survive in Britain?" (PDF). Ecological Research Centre Faunaconsult, Tegelseweg. Retrieved 5 January 2016.
  18. ^ a b c Cieslak, Michael; Pruvost, Melanie; Benecke, Norbert; Hofreiter, Michael; Morales, Arturo; et al. (2010). "Origin and History of Mitochondrial DNA Lineages in Domestic Horses". PLOS ONE. 5 (12:e15311): e15311. Bibcode:2010PLoSO...515311C. doi:10.1371/journal.pone.0015311. PMC 3004868. PMID 21187961.
  19. ^ a b c Lindgren, Gabriella; Backstrom, Niclas; Swinburne, June; Hellborg, Linda; Einarsson, Annika; et al. (1985). "Limited number of patrilines in horse domestication". Nature Genetics. 36 (4): 335–6. doi:10.1038/ng1326. PMID 15034578. S2CID 38310144.
  20. ^ Stuart, Tony (2006)"Exotic world before Suffolk", British Archaeology (86). Retrieved 22 March 2011. Archived page Archived 19 July 2012 at the Wayback Machine
  21. ^ Daniel 1950, p. 173.
  22. ^ Ling, Yinghui; Ma, Yuehui; Guan, Weijun; Cheng, Yuejiao; Wang, Y.; et al. (2010). "Identification of Y chromosome genetic variations in Chinese indigenous horse breeds". Journal of Heredity. 101 (5): 639–43. doi:10.1093/jhered/esq047. PMID 20497969.
  23. ^ Jansen, Thomas; Forster, Peter; Levine, Marsha A.; Oelke, Hardy; Hurles, Matthew (2002). et al. "Mitochondrial DNA and the origins of the domestic horse". PNAS. 99 (16): 10905–10910. Bibcode:2002PNAS...9910905J. doi:10.1073/pnas.152330099. PMC 125071. PMID 12130666.
  24. ^ Vilà, Carles; Leonard, Jennifer A.; Götherström, Anders; Marklund, Stefan; Sandberg, Kaj; et al. (2001). "Widespread origins of domestic horse lineages". Science Magazine. 291 (5503): 474–77. Bibcode:2001Sci...291..474V. doi:10.1126/science.291.5503.474. PMID 11161199.
  25. ^ a b McGahern, A. M.; Edwards, C. J.; Bower, M. A.; Heffernan, A.; Park, S. D. E.; et al. (2006). "Mitochondrial DNA sequence diversity in extant Irish horse populations and in ancient horses". Animal Genetics. 37 (5): 498–502. doi:10.1111/j.1365-2052.2006.01506.x. PMID 16978181.
  26. ^ Woodman, Peter; McCarthy, Margaret; Monaghan, Nigel (1997). "The Irish quaternary fauna project". Quaternary Science Reviews. 16 (2): 129–59. Bibcode:1997QSRv...16..129W. doi:10.1016/S0277-3791(96)00037-6.
  27. ^ Jordana, J; Pares, P. M; Sanchez, A (1995). "Analysis of genetic relationships in horse breeds" (PDF). Journal of Equine Veterinary Science. 15 (7): 320–328. doi:10.1016/S0737-0806(06)81738-7. Archived from the original (PDF) on 3 March 2016. Retrieved 23 April 2012.
  28. ^ Johns & Potter 2002, p. 24.
  29. ^ Brown, Bennett & Rhodes 2009, pp. 50–61.
  30. ^ Johns & Potter 2002, p. 28.
  31. ^ Budd 1998.
  32. ^ "Exmoor Pony". Exmoor Ponies in Conservation. Archived from the original on 1 October 2011. Retrieved 17 April 2011.
  33. ^ Grout, James. "The British War-Chariot". Encyclopedia Romana. University of Chicago. Retrieved 29 September 2015.
  34. ^ Johnstone, Cluny Jane. "A Biometric Study of Equids in the Roman World" (PDF). University of York. Archived from the original (PDF) on 5 October 2012. Retrieved 17 April 2011.
  35. ^ Wynmalen, Henry (1971). Horse Breeding & Stud Management. J. A. Allen. p. 83. ISBN 978-0-85131-139-5.
  36. ^ "Exmoor pony Bear is top of the world". West Somerset Free Press. 6 January 2012. Retrieved 24 February 2016.
  37. ^ "Bear's on top of the world again". West Somerset Free Press. 3 January 2013.
  38. ^ "Czechs import wild horses to save biodiversity". cbsnews.com.
  39. ^ "Wild horses and aurochs on pasture are now streaming". idnes.cz.

Bibliography

Further reading

  • Morrison, Alex (1980). Early man in Britain and Ireland: an introduction to Palaeolithic and Mesolithic cultures. Croom Helm. ISBN 978-0-85664-084-1.
  • Smith, Christopher (1992). Late Stone Age hunters of the British Isles. Routledge. ISBN 978-0-415-07202-1.

External links

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